Key to the Juglandaceae of Iowa

A. Pith of twigs chambered, interrupted by crosswalls; nut with a distinctly furrowed shell, husk not splitting into sections, husk rotting away from nut with age after fruit-fall. Juglans
Butternut & Walnut
A. Pith of twigs solid, crosswalls lacking; shell of nut with a ± smooth surface, husk splitting into four sections at maturity. Carya
Hickory
Carya: Hickories
1. Terminal bud scales valvate, not over-lapping [2 scales visible, meeting at edges & essentially not overlapping]. [2]
1. Terminal bud scales imbricate [> 2 scales visible & overlapping]. [3]
2. Mature leaves commonly with ≤ 9 leaflets; buds yellow; nut essentially spherical,
husk splitting to approximately the middle of the nut.
C. cordiformis
Bitternut Hickory
2. Mature leaves commonly with ≥ 11 leaflets; buds brownish; nut cylindrical,
husk splitting to the base of the nut.
C. illinoinensis
Pecan
3. Leaves primarily with 5 [or fewer] leaftlets. [4]
3. Leaves primarily with 7 [or more] leaftlets. [5]
4. Teeth of leaflets bearing tuft of pubscence just beneath tooth apex; nut usually > 3.5 cm; husk of fruit thick [typically ≥ 0.5 cm], splitting to base of fruit at maturity; mature bark shaggy, splitting vertically into long loose plates; widespread in the state. Carya ovata
Shagbark Hickory
4. Teeth of leaftlets lacking tuft of pubescence; nut usually < 3.5 cm; husk of fruit thin [typically ≤ 0.5 cm], splitting to approximately the middle of the fruit; mature bark tight, becoming deeply furrowed with age, not shaggy; reported from SE Iowa [Van Buren county]. Carya glabra
Pignut Hickory
5. Mature bark shaggy, splitting vertically into long loose plates; leaves ≥ 60 cm long; fruits typically ≥ 5 cm long; twigs stout. Carya laciniosa
Shellbark Hickory
5. Bark ridged to deeply furrowed, not shaggy; leaves ≤ 60 cm long; fruits typically ≤ 5 cm long; twigs stout or slender. [6]
6. Leaves typically bearing ≥ 7 leaflets; lower leaf surface, petiole, and leaf rachis ± densely pubescent; husk of nut splitting to base of fruit; twigs stout; reported from SE & SW corners of Iowa. Carya tomentosa
Mockernut Hickory
6. Leaves typically bearing ≤ 7 leaflets; lower leaf surface, petiole, and leaf rachis essentially glabrous; husk of nut splitting to middle of fruit; twigs slender; reported from SE Iowa [Van Buren county]. Carya glabra
Pignut Hickory
Juglans: Butternut & Walnut
7. Pith of twigs chocolate brown; nut elliptical, surface sticky-hairy; leaf scar with linear moustache-like upper edge. Juglans cinerea
Butternut
7. Pith of twigs tan; nut spherical, the surface smooth; leaf scar lacking linear moustache-like upper edge. Juglans nigra
Black Walnut

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